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Online ISSN:
2466-4367

Issue 21, (2026)

Published:
22.01.2026.

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Reforesta is an Open Access, peer-reviewed online journal published by the Scientific and Professional Society Reforesta.

Reforesta publishes original papers on the fundamental and applied aspects of tree improvement, plant production and quality, and afforestation/reforestation.

Reforesta will consider only original manuscripts that have not been published elsewhere and are not under consideration by another journal.

No article processing charge is applied.

01.12.2017.

Review paper

Seedling Establishment on a Forest Restoration Site

Seedling field performance is affected by both their quality and reforestation site conditions. Seedlings enter the establishment phase when they start to develop root systems into the surrounding soil and are coupled to the restoration site. Once seedlings are established, their inherent growth potential is related to morphological and physiological attributes and their ecophysiological response to site environmental conditions, which ultimately determines field performance. This establishment phase is a time when seedlings developed with certain nursery cultural practices begin to respond to site conditions. This phase is also a period when silvicultural practices have created microsites intended to benefit established seedlings field performance. Seedlings can be exposed to a wide range of environmental conditions during the establishment phase, some of which may be extreme enough to exceed their ability to physiologically tolerate environmental stress. When this occurs, seedling growth on the restoration site is reduced. On the other hand, this phase can provide planted seedlings with ideal environmental conditions that allow for an optimum physiological response and maximization of their growth potential. An understanding of the ecophysiological capability of planted seedlings can ensure they have the best chance at rapid stand establishment.

Steven Grossnickle

01.12.2021.

Research paper

Net benefits of silky oak (Grevillea robusta) for small farmers in Musanze District, Rwanda

This study was conducted in Musanze district, Rwanda, to evaluate the net benefits of silky oak (Grevillea robusta) production for small farmers. A semi-structured questionnaire was administered to 100 households distributed in four villages. The cost-benefit ratio was used as a decision-making tool. A diameter tape and a Haga altimeter were used to measure the DBH and height of grevillea and other dominant agroforestry tree species. Results indicate that 66 % of laborers were family members, gaining 625 Rwf per person day with grevillea production (US$ 0.61, with 4-8 hours of work per day. Also, results show that farmers earn 57,950 Rwandan Francs (Rwf) per hectare per year (US$ 57.48 from grevillea products (stakes, poles, charcoal, and timber), whereas the mean investment in grevillea production is 54,200 Rwf ha-1 year-1 (US$ 53.76. The net farm income is 3,225 Rwf (US$ 3.2 ha-1 year-1. The net benefit from grevillea product is affected by poor farmer data record as the majority of67% of respondents has not attended any formal education. However, farmers gain additional benefits of US$ 628, US$ 298, and US$ 224ha-1 year-1 from potatoes, maize, and bean, respectively, depending on their crop choice during intercropping with grevillea. The benefit-cost ratio is 1.06, which highlights the modest profitability of growing grevillea on the farm. In addition, soil erosion control, soil fertility increase, landslide prevention, shade provision, microclimate improvement, and biodiversity conservation were reported as ecosystem services of grevillea on the farm. Growth performance of grevillea (diameter at breast height, tree height, and volume) is analyzed in all four villages. The study shows that growing grevillea is profitable for smallholder farmers via tree products and farm benefits, including ecosystem services aspects.

Rene Niyomfura, Gerald Kapp, Canisius Patrick Mugunga, Jean de Dieu Niyomugabo

01.12.2019.

Research paper

Growth and quality of Pinus nigra (Arn.), Pinus sylvestris (L.) and Pinus pinaster (Aiton) seedlings in two container types

Abstract In this study we investigated the dynamics of growth and quality features of 1+0 Austrian Pine (AP; Pinus nigra Arn.), Scots Pine (SP; Pinus sylvestris L.)  and Maritime Pine (MP; Pinus pinaster Aiton) seedlings in containers Yukosad (YS; hard plastic, 75cm3, 610 seedlings m-2) and Siset (SS; gray cardboard with white coating, 128 cm3, 589 seedlings m-2). The experiment, for all Pine species, was conducted in randomized blocks in four replications. Each replication contained 10 multipots of each container type. After the seed germination, every 15 days, i.e. 10 times during the vegetation season, 10 seedlings of each of 4 replications, i.e. 40 seedlings of each container and Pine species, were analyzed. The following morphological characteristics were measured: shoot height (SH), root collar diameter (RCD), shoot dry weight (SDW), number, length and dry weight (RDW) of I, II and III order lateral roots (FOLR, SOLR and TOLR) and of the central root. Quality ratios and indexes were calculated. The dynamics of growth of selected features during the vegetation season was expressed as trend lines. AP and SP seedlings, grown in containers YS had higher values of almost all measured traits (SH, RCD, SDW, number, length and dry weight of FOLR, SOLR and TOLR). However, in AP, the SS seedlings had better values of quality indexes and ratios (SQ, DQI, SDW/RDW and RDW/SDW ratio), than YS seedlings. SP seedlings, except for SQ, had better quality indexes in YS container. MP seedlings, with some exceptions, have better above ground quality features in SS containers, while root systems don’t differ significantly in both containers. Root quality indicators (root intensity and root bound indexes) are more favorable in SS containers in all Pine species. The polynomial trend lines in AP and SP grow in similar way and intensity in both containers, gradually increasing during the vegetation season. MP seedlings differ in much more intensive growth of above ground features, especially in the half of the vegetative period in both containers, while during this period root quantity and length stagnates. Generally, the quality of AP and SP seedlings is more or less equal and satisfactory in both containers, while MP seedlings show slightly better results in SS container.

Dana Dina Kolevska, Anastazija Dimitrova, Kristijan Cokoski, Mare Basova

01.12.2024.

Research paper

The bigger the tree the better the seed – effect of Sessile oak tree diameter on acorn size, insect predation, and germination

Sessile oak is one of the ecologically and economically most important tree species in Europe. Recently, the importance of this species has been particularly highlighted in the context of climate change, where it is expected that these species will play a significant role within their natural range and beyond. To regenerate and expand the range of the sessile oak forests, a large amount of healthy acorns is needed. The acorn production and the acorns themselves are endangered by a large number of abiotic and biotic factors. As oak trees of different sizes do not produce the same amount of seed of the same quality, we investigated how the sessile oak tree diameter affects the acorn size, insect infestation, and germination rate. On the other side, as the acorn size influences the attack rates of the most significant acorn pest – Curculio glandium (Marsham, 1802), and the germination rates of both the damaged and healthy acorns we also investigated how the acorn dimensions influence the insect damage and germination rates, and how the insect damage influences the germination rates. We determined that the oak tree size influences some oak seed characteristics that are important for forest regeneration, while it does not affect others. A greater acorn yield was recorded on the bigger oak trees. The tree dimensions also had a significant influence on the acorn size. Greater average length, as well as greater average acorn diameter, were recorded on bigger trees. Although the tree size affects the acorn size, it does not influence the acorn insect predation rates or its germination rates. These parameters are affected by the acorn size itself. Bigger acorns had a greater germination rate. Insects preferred smaller acorns and had a significantly negative influence on the germination rate.

Jovan Dobrosavljevic, Jovana Devetaković, Branko Kanjevac

01.12.2024.

Research paper

Testing the reliability of morphological patterns to identify Sonderegger pine in forest tree seedling nurseries

Sonderegger pine (Pinus x sondereggeri H.H. Chapm.), the natural hybrid of longleaf (Pinus palustris Mill.) and loblolly pine (Pinus taeda L.), commonly occurs in longleaf pine seedlots grown in forest tree seedling nurseries in the southeastern United States. Because longleaf pine seedlings have a grass stage with minimal epicotyl development (< 1 cm), the initiation of stem growth (12 to 15 cm) in longleaf pine seedlots has been used to indicate that hybridization with loblolly pine (Pinus taeda L.) has occurred. Sonderegger pine seedlings are typically culled at the nursery due to observations of poor form and wood quality after outplanting. However, research documenting Sonderegger pine seedling morphology has not been published for more than 60 years, and to our knowledge, no seedling quality assessments have been made. To better understand how morphological traits of longleaf and loblolly pine are expressed in hybrid seedlings, stem length, hypocotyl length, and root collar diameter (RCD) were compared among one-year-old container-grown longleaf, loblolly, and seedlings visually classified as Sonderegger pine. Sonderegger pine seedlings had a range of stem development, with most (62%) seedlings measuring < 12 cm tall. Some pure longleaf pine seedlings had up to 10 cm of stem elongation, but the cause of early height growth in these seedlings is unknown. More than 90% of Sonderegger pine seedlings met or exceeded RCD recommendations for planting loblolly (≥ 3.2 mm) and longleaf pine (≥ 4.75 mm).

Kelsey Shoemaker, D. Paul Jackson, Joshua Adams

01.12.2016.

Review paper

Reforestation in Macedonia: History, current practice and future perspectives

This paper presents data about forestry and reforestation in the Republic of Macedonia. The country is characterized with great diversity of natural conditions and rich floral and faunal biodiversity Forests in Macedonia cover 38% of its territory. About 71% represent coppiced and degraded and 29% tall forests. Historical, social and natural conditions caused gradual deforestation, forest and land degradation. Foundations of artificial afforestation were laid in the first decade of XX century. First reforestation started already in 1913/14 and continued, with various intensity, in the next decades. In the period between two world wars a foundation of modern forestry was established, as forestry education, scientific work etc., to help dealing with reforestation of waste bare and erosive lands. The most intensive reforestation was performed in 1971-1990 and during the following years significantly decreases. There is room for improving of some aspects of the reforestation, in aim to improve survival and development of the young stands.

Dana Dina Kolevska, Ivan Blinkov, Pande Trajkov, Vladimir Maletić

01.12.2016.

Research paper

Using double-sampling techniques to reduce the number of measurement trees during forest inventories

Variable-radius sampling techniques are commonly used during forest inventories. For each sample tree at a particular sampling point, diameter and height(s) are measured and then weight is estimated using established equations.  Heights can require a fair amount of time to measure in the field.  Separating the weight per acre estimate into two components; average basal area per acre and WBAR (individual tree weight-basal area ratio) across all points, can often lead to more efficient sampling schemes. Variable-radius sampling allows for a quick estimate of basal area per acre at a point since no individual tree measurements are needed.  If there is a strong relationship between weight and basal area, then by knowing basal area you essentially know weight.  Separation into two components is advantageous because in most cases there is more variability among basal area estimates per point then there is in WBAR. Hence, you can spend more resources establishing many points that only estimate basal area – often called “Count” points. “Full” points are those where individual tree measurements are also conducted. There is little published information quantifying the impacts on basal area, weight, etc., estimates among different “Full/Count” sample size ratios at the same site. Inventories were examined to determine this method’s applicability to loblolly pine plantations in southern Arkansas and northern Louisiana. Results show there is more variability among basal area estimates than WBAR and that the amount of trees being “intensively” measured is excessive.  Based on these four plantations, a “Full” point could be installed ranging from every other point to every fifth point depending on site conditions and the desired variable.

Curtis L. VanderSchaaf, Gordon Holley, Joshua Adams

01.12.2024.

Research paper

Morphological assessment of plus trees progeny as a basis for establishing a sessile oak seed orchard

In the context of variable climate conditions, selecting high-quality Sessile oak (Quercus petraea (Matt.) Liebl.) reproductive material is crucial for sustainable forestry. This research aimed to assess the genetic potential of 86 selected plus trees by analyzing seedling morphological characteristics and quality indices. The ultimate goal was to identify the most promising genotypes for establishing seed orchards. Root collar diameter (D), seedling height (H), stem dry mass (SDM), and root dry mass (RDM) were measured under uniform nursery conditions. The following derived indices were calculated: seedling height to root collar diameter ratio (H/D), stem dry mass to root dry mass ratio (SDM/RDM), and Dickson’s quality index (DQI). Descriptive statistics, one-factorial analysis of variance (ANOVA) and Pearson correlation analysis were performed as well. The results showed significant variability in all observed morphological characteristics and quality indices among half-sib families, confirming a strong genetic influence. Correlation analysis indicated strong positive relationships between DQI and all primary morphological characteristics (r≤0.94), confirming DQI as the most effective integrative indicator of seedling quality. The combined ranking of the half-sib families, based on DQI, H/D, and SDM/RDM, identified those with optimal morphological characteristics, including an optimal H/D ratio, balanced SDM/RDM, and a high DQI value. The identified superior half-sib families represent valuable genetic material for the establishment of future seed orchards, thereby contributing to the production of better-quality seedling material and to improved forest ecosystem resilience under climate change.

Vladan Popović, Sanja Lazić, Aleksandar Lučić, Ljubinko Rakonjac, Vanja Daničić, Boris Ivanović, Aleksandra Petrović

01.12.2017.

Review paper

Cost of inoculating seedlings with Pisolithus tinctorius spores

Although the production of commercial products of vegetative Pt (Pisolithus tinctorius (Pers.) Coker & Couch) inoculums has ceased in North America due to a lack of demand by consumers, the number of products that contain Pt spores has increased.  The quality, quantity and price of these products vary considerably.  The cost of inoculating 1,000 tree seedlings with Pt basidiospores can vary from $0.45 to more than $30.  The cost of treating with Pt spores is lowest when seedlings are inoculated in a container nursery using rates that are less than 0.4 mg per seedling.  However, with some products the cost to treat 1,000 bareroot seedlings is greater than $500 when spores are applied in the planting hole.  Three decades ago, 1 g of Pt spores could be purchased for $0.13 and now the price of 1 g can exceed $14. Although many research papers provide data on the biological response to inoculating seedlings with spores, few document the cost of inoculation. Commercial products that are marketed toward homeowners containing both ectomycorrhizal and endomycorrhizal spores are more expensive than products that contain only ectomycorrhizal spores.  In situations where survival and growth of seedlings are not increased, the benefit/cost ratio will typically be less than one.

David B. South